Good find! Honestly, the DBV would have been my last guess. Glad you didn't give up on just the usual suspects. But as I've found with these machines, and you now have too, something completely unrelated can cause errors that point you in the total wrong direction.
When I had Winning Streak I was having problems with it locking up and the Dotmation displaying 'Bad Packet' or 'Bad Page' with like an '0XFF0011', something like that. Totally random times and out of nowhere. I was focused on the Dot system, board, power supply, chips. I began to think it was a bad Rom or something, but when I was doing housekeeping on it that's when I noticed a piece of random scotch tape under the I/O board in the bottom cage. It was making weak transient connections between incorrect contacts, screwing things up. Removed the tape, never had the problem again. That's why I mentioned tape in the previous post. It seems so innocuous, and people would think 'ehh, that won't make a difference.', but it can. If it's not supposed to be there, it doesn't belong there.
As these machines age, I'm sure this won't be your only headscratcher. But you have the right mindset for troubleshooting, that's important! Know that these machines themselves are strong (except for the power supplies) and things don't often 'officially' break on their own.