Just to update this thread, everything is working again, but what I did doesn’t really explain it.
All I did was unplug the green cable from signature card to bypass on the front cpu door. Boot it, it came up fine. Turned it off, plugged it back into signature card, booted. IGT Signature Not Found! Again.. power cycled one more time and it’s working flawlessly now.
Maybe the cable wasn’t seated properly, but who knows.